The rituals displayed can convey reproductive fitness or they can just trick reluctant mates into copulation.

所展示的仪式可以传达生殖能力，也可以只是欺骗不情愿的配偶进行交配。

Particularly when they are engaged in other activities, like copulation, which requires coming partway out of the shell.

尤其是当它们忙于其他活动时，比如交配，这需要它们的部分身体从壳里出来。”

(Laughter) Now, we define extra-pair copulation as the mating outside of a pair bond.

（笑声） 现在，我们将配对之外的交配定义为配对之外的交配。

She said there was forced copulation the whole time she was here.

她说在她工作期间 一直存在强迫性行为。

The forced copulation? - We don't know that.

强迫性交的事 -还不确定。

For males, it might just be a few moments of copulation and that's it.

对于雄性来说，这可能只是短暂的交配，仅此而已。

CHRISTINE WILKINSON: These lezzy lizzies actually simulate copulation with each other, which increases their fertility.

克里斯汀·威尔金森：这些女同性恋者实际上模拟了彼此的交配，这提高了她们的生育能力。

Researchers tested to see whether showing several demonstrations of copulation would lead to a higher percentage of learned mate preference.

研究人员进行了测试，以了解展示多次交配示范是否会导致更高比例的习得择偶偏好。

Longer bacula can help a male compete, because they're associated with what's called prolonged intromission, or longer periods of copulation.

更长的杆状骨可以帮助雄性竞争，因为它们与所谓的长时间插入或更长的交配时间有关。

But is there anything above and beyond that that is the result of actual copulation and interacting with those spines?

但除此之外还有什么是实际交配和与这些刺相互作用的结果吗？