Price equation

In the theory of evolution and natural selection, the Price equation (also known as Price's equation or Price's theorem) describes how a trait or gene changes in frequency over time. The equation uses a covariance between a trait and fitness to give a mathematical description of evolution and natural selection. It provides a way to understand the effects that gene transmission and natural selection have on the proportion of genes within each new generation of a population. The Price equation was derived by George R. Price, working in London to re-derive W.D. Hamilton's work on kin selection. The Price equation also has applications in economics. Examples of the Price equation can be found here: Price equation examples.