The researchers were from the University of ColoradoDenver, Denver, Colorado; Washington University, St. Louis, Missouri; and the University of Pennsylvania, Philadelphia, Pennsylvania.
研究人员来自科罗拉多丹佛大学，华盛顿大学，圣路易斯大学，密苏里大学及宾夕法尼亚，费城的大学。
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Even the tracks alone say a lot about the feeding behavior of the ancient birds, commented paleontologist Martin Lockley of the University of Colorado in Denver.
丹佛科罗拉多大学的古生物学家马丁?劳克雷评论说，即使只有这些足迹也很能说明这些古代鸟类的进食行为。
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"I don't want to cry Wolf on this," said Dr. John M. Kittelson, a professor in the Department of Biostatistics and Informatics at the University of ColoradoDenver.