From the position of the legs, the curve of the abdomen, and the lack of wing marks, the researchers suspect that the imprint was made by an ancient mayfly that held its wings upright when at rest.

The formation point-roils were difficult, for through the inverted half Jonathan had to think upside down, reversing the curve of his wing, and reversing it exactly in harmony with his instructor's.

An especial perturbation technique of curve fitting shape function is applied for variation of the wing shape.