However, the future approaches for GWAS should be based on the multi-locus genetic model, QTN-by-environment interaction, epistatic detection and multivariate analysis.
但是， 今后的方法应当是以多位点模型、环境互作、上位性检验和多个相关性状联合分析为主。
3
The interaction effects influenced the direct effects. 3. The drought tolerance related traits were determined not only by both additive QTLs and epistatic QTLs.